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HyPer has two excitation maxima at 420 nm and 500 nm corresponding to the protonated and anionic form of YFP respectively, and one emission peak at 516 nm.(817) Oxidation of HyPer by H 2 O 2 results in a decrease in 420 nm excitation peak and a proportional increase in the 500 nm excitation peak, permitting a ratiometric readout.(818) roGFP2-based sensors also have similar spectroscopic properties to HyPer (excitation maxima at 405 nm (reduced) and 488 nm (oxidized) and emission maximum at 510 nm).(814) Both methods have several additional benefits over most small molecule probes