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However, in contrast to some other invertebrate groups with anhydrobiosis where trehalose levels in the dry state reach 15–20% of the dry weight (nematodes, Artemia , and the sleeping chironomid Polypedilum vanderplanki ), tardigrades have much lower levels of trehalose (below 3%), and only in one family (Macrobiotidae) is its level upregulated in response to desiccation [82,83], while many tardigrade species produce low or undetectable levels of the disaccharide [49,82,83,84,85]